Late Pleistocene brown bear (Ursus arctos) from the Caucasus

Baryshnikov G.F.

P. 9-17

The examination of the fossil collections of teeth and bones of Ursus arctos from Paleolithic sites in Kudaro cave, South Ossetia, revealed the attribution of the Late Pleistocene material to the fossil subspecies U. a. binagadensis. The brown bear remains recovered in the layers dated by the end of Late Pleistocene is referred to the extant Caucasian subspecies U. a. meridionalis.DOI: 10.15298/rusjtheriol.9.1.02

• Adlerberg G.P. 1935. [Bears of Caucasus. Systematic review] // Izvestiya Akademii Nauk SSSR. Otdelenie matematicheskikh i estestvennykh nauk. No.1. P.73-100 [in Russian].
• Arabuli A.B. 1986. [About intraspecific variation of brown bear of Georgia] // Izvestiya Akademii Nauk GSSR. Seriya Biologicheskaya. T.12. No.6. P.409-415 [in Russian].
• Baryshnikov G. 1999. Bone assemblages from Acheulean and Mousterian levels in the Kudaro Caves in the Caucasus Mountains // The Role of Early Humans in the Accumulation of European Lower and Middle Palaeolithic Bone Assemblages. Monographien des romisch-germanischen Zentralmuseums, Mainz. Bd.42. P.237-253.
• Baryshnikov G.F. 2002. Local biochronology of Middle and Late Pleistocene mammals from the Caucasus // Russian Journal of Theriology. Vol.1. No.1. P.61-67.
• Baryshnikov G.F. 2007. [Bears (Family Ursidae)]. Sankt-Peterburg: Nauka. 542 p. [in Russian].
• Baryshnikov G.F. 2010a. Middle Pleistocene Ursus thibetanus (Mammalia, Carnivora) from Kudaro caves in the Caucasus // Proceeding of the Zoological Institute RAS. Vol.314. No.1. P.67-79.
• Baryshnikov G.F. 2010b. Pleistocene Mustelidae (Carnivora) from Paleolithic site in Kudaro Caves in the Caucasus // Russian Journal of Theriology. Vol.8. No.2. P.75-95.
• Bobrinsky N.A. Kuznetsov B.A. & Kuzyakin A.P. 1944. [Key for Mammals of USSR]. Moskva: Sovetskaya Nauka. 440 p. [in Russian].
• Driesch A., von den. 1976. A guide to the measurement of animal bones from archaeological sites // Peabody Museum Bulletin. No.1. P.1-136.
• Ellerman J.R. & Morrison-Scott T.C.S. 1951. Checklist of Palaearctic and Indian Mammals, 1758 to 1946. London: British Museum (Natural History). 810 p.
• Chestin I. & Mikeshina N. 1998. Variation in skull morphology of brown bears (Ursus arctos) from Caucasus // Journal of Mammalogy. Vol.79. No.1. P.118-130.
• Gromova V. 1960. [Determination key to mammals of USSR based on postcranial bones. Part 2. Determination key based on large tarsal bones]. Moskva: Izdatel'stvo Akademii Nauk SSSR. 118 p. [in Russian].
• Heptner V.G., Naumov N.P., Yurgenson P.B., Sludsky A.A., Chirkova A.F. & Bannikov A.G. 1967. [Mammals of Soviet Union. T.2. Parts 1. Sea Cows and Carnivores]. Moskva: Vysshaya Shkola. 1004 p. [in Russian].
• Lobachev V.S., Chestin I.E., Kudaktin A.N. & Fomin S.V. 1988. [Character of territorial utilization of bears from different ecomorph in Western Caucasus] // Byulleten' Moskovskogo Obshchestva Ispytatelei Prirody. Otdelenie Biologicheskoe. T.93. No.1. P.23-33 [in Russian].
• Mezhlumian S.K. 1959. [Bear from beach settlements of Sevan Lake] // Izvestiya Akademii Nauk Armyanskoi SSR. Vol.12. No.4. P.85-88 [in Russian].
• Murtskhvaladze M., Gabashelishvili A. & Tarkhnishvili D. 2010. Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus // Molecular Ecology. Vol.19. No.9. P.1829-1841.
• Novikov G.A. 1956. [Carnivorous Mammals of USSR Fauna]. Moskva-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 295 p. [in Russian].
• Ognev S.I. 1931. [Mammals of Eastern Europe and Northern Asia]. T.2. Moskva-Leningrad: Glavnauka. 776 p. [in Russian].
• Vekua A.K. 1978. [Fossil vertebrates from Tsutskhvati caves] // (Davitaya F.F., ed.) Izuchenie Peshcher Kolkhidy. Tsutskhvatskaya Mnogoyarusnaya Karstovaya Peshchernaya Sistema. Tbilisi: Metsniereba. P.94-126 [in Russian].
• Vereshchagin N.K. 1947. [New race of brown bear from Binagady asphalt (Ursus arctos binagadensis, susp. nova) // Doklady Akademii nauk Azerbaidzhanskoi SSR. T.2. No.8. P.346-348 [in Russian].
• Vereshchagin N.K. 1959. [Mammals of Caucasus. History of Fauna Origin]. Moskva-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 704 p. [in Russian].